Ownership Submission
FORM 4
Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).
UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP OF SECURITIES

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public Utility Holding Company Act of 1935 or Section 30(h) of the Investment Company Act of 1940
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(Print or Type Responses)
1. Name and Address of Reporting Person *
LACOB JOSEPH
  2. Issuer Name and Ticker or Trading Symbol
ALIGN TECHNOLOGY INC [ALGN]
5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)
__X__ Director _____ 10% Owner
_____ Officer (give title below) _____ Other (specify below)
(Last)
(First)
(Middle)
C/O ALIGN TECHNOLOGY INC., 881 MARTIN AVE
3. Date of Earliest Transaction (Month/Day/Year)
03/27/2008
(Street)

SANTA CLARA, CA 95050
4. If Amendment, Date Original Filed(Month/Day/Year)
6. Individual or Joint/Group Filing(Check Applicable Line)
_X_ Form filed by One Reporting Person
___ Form filed by More than One Reporting Person
(City)
(State)
(Zip)
Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1.Title of Security
(Instr. 3)
2. Transaction Date (Month/Day/Year) 2A. Deemed Execution Date, if any (Month/Day/Year) 3. Transaction Code
(Instr. 8)
4. Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4 and 5)
5. Amount of Securities Beneficially Owned Following Reported Transaction(s)
(Instr. 3 and 4)
6. Ownership Form: Direct (D) or Indirect (I)
(Instr. 4)
7. Nature of Indirect Beneficial Ownership
(Instr. 4)
Code V Amount (A) or (D) Price
Common Stock 03/27/2008 03/27/2008 M   60,000 A $ 0.05 758,327 (1) D  
Common Stock 03/27/2008 03/27/2008 M   8,000 A $ 9.8 766,327 D  
Common Stock 03/27/2008 03/27/2008 M   8,000 A $ 4.04 774,327 D  
Common Stock 03/27/2008 03/27/2008 M   43,000 A $ 6.15 817,327 D  
Common Stock 03/27/2008 03/27/2008 M   8,000 A $ 8.84 825,327 D  
Common Stock 03/27/2008 03/27/2008 M   8,000 A $ 7.19 833,327 D  
Common Stock 03/27/2008 03/27/2008 M   8,000 A $ 7.35 841,327 D  
Common Stock               364,006 I Trust for reporting person
Common Stock               148,767 I Trust for children
Common Stock               746,210 (2) I By KCPB VIII
Common Stock               43,263 (2) I By KPCB VIII FF
Common Stock               20,258 (2) I By KPCB Life

Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
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Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned
(e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security
(Instr. 3)
2. Conversion or Exercise Price of Derivative Security 3. Transaction Date (Month/Day/Year) 3A. Deemed Execution Date, if any (Month/Day/Year) 4. Transaction Code
(Instr. 8)
5. Number of Derivative Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4, and 5)
6. Date Exercisable and Expiration Date
(Month/Day/Year)
7. Title and Amount of Underlying Securities
(Instr. 3 and 4)
8. Price of Derivative Security
(Instr. 5)
9. Number of Derivative Securities Beneficially Owned Following Reported Transaction(s)
(Instr. 4)
10. Ownership Form of Derivative Security: Direct (D) or Indirect (I)
(Instr. 4)
11. Nature of Indirect Beneficial Ownership
(Instr. 4)
Code V (A) (D) Date Exercisable Expiration Date Title Amount or Number of Shares
Stock Option (right to buy) $ 0.05 03/27/2008 03/27/2008 M     60,000 05/29/1999 05/29/2008 Common Stock 60,000 $ 0 0 D  
Stock Option (right to buy) $ 9.8 03/27/2008 03/27/2008 M     8,000 05/15/2002 05/15/2011 Common Stock 8,000 $ 0 0 D  
Stock Option (right to buy) $ 4.04 03/27/2008 03/27/2008 M     8,000 05/16/2003 05/16/2012 Common Stock 8,000 $ 0 0 D  
Stock Option (right to buy) $ 6.15 03/27/2008 03/27/2008 M     43,000 04/23/2004 04/23/2013 Common Stock 43,000 $ 0 0 D  
Stock Option (right to buy) $ 8.84 03/27/2008 03/27/2008 M     8,000 05/15/2004 05/15/2013 Common Stock 8,000 $ 0 0 D  
Stock Option (right to buy) $ 7.19 03/27/2008 03/27/2008 M     8,000 05/25/2006 05/25/2015 Common Stock 8,000 $ 0 0 D  
Stock Option (right to buy) $ 7.35 03/27/2008 03/27/2008 M     8,000 05/24/2007 05/24/2016 Common Stock 8,000 $ 0 0 D  

Reporting Owners

Reporting Owner Name / Address Relationships
 Director  10% Owner  Officer  Other
LACOB JOSEPH
C/O ALIGN TECHNOLOGY INC.
881 MARTIN AVE
SANTA CLARA, CA 95050
  X      

Signatures

 Roger E. George, Atty-in-Fact for Joseph S. Lacob   03/28/2008
**Signature of Reporting Person Date

Explanation of Responses:

* If the form is filed by more than one reporting person, see Instruction 4(b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
(1) Since the date of the reporting person's last ownership report, and in connection with acquisition of shares reported on this Form 4, he transferred an aggregate of 788,650 shares of Common Stock pursuant to a divorce settlement. The reporting person no longer reports as beneficially owned any shares held by his ex-wife.
(2) Mr. Lacob is a general partner of KPCB VIII Associates, L.P., a CA limited partnership ("KPCB VIII Associates"). KPCB VIII Associates is the general partner of Kleiner Perkins Caufield & Byers VIII, L.P., a CA limited partnership ("KPCB VIII") and KPCB VIII Founders Fund L.P., a CA limited partnership ("KPCB VIII FF"). Mr. Lacob is also a general partner of KPCB VII Associates, L.P., a CA limited partnership ("KPCB VII Associates"). KPCB VII Associates is the general partner of KPCB Life Sciences Zaibatsu Fund II, L.P., a CA limited partnership ("KPCB Life"). Mr. Lacob disclaims beneficial ownership of the shares of the Issuer's common stock held directly by KPCB VIII, KPCB VIII FF, KPCB Life, except to the extent of any indirect pecuniary interest in his distributive share therein.

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